The Great Dane
First Grade
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It depends how how you are defining biggest.The 3 biggest nations in RU have traditionally been Australia, NZ and SA
Apparently if arguably the 2 biggest RU nations fall off it means nothing...
BTW, Australia hasn't been shit in RU for the better part of 20 years now, and if anything that's benefited WR by making the WC more competitive. The All Blacks dominance is already tailing off, and it's only benefited WR so far as well.